This book aims to encourage the reading of "On the Origin of Species" and to include it in the teaching of evolution. With a comprehensive overview of the development of Darwins theory, the volume provides relevant aspects of Darwins life and work in connection with the broader context of his time. The historical and philosophical analysis, mirrored in the socio-cultural scope, enables the diachronic reading of the text. It is built on various sources of historians and philosophers of science and sheds fresh light on them. Its uniqueness is the broad structure that covers four parts: the pre-Darwinian concepts of species changes; some key elements of Darwins pursuit of the causes of evolution, from his voyage on Beagle to the publication of his groundbreaking work; chapter-by-chapter analysis of the "Origin"; and subsequent developments in evolutionary thought. This book is of interest to undergraduate and graduate students, scholars in history, philosophy, and sociology of science and science education, as well as the general public.